Fly Anatomy (FA): adult head

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Phenotype Ontology
Like Gene Ontology (GO), phenotypy ontology classifies and organizes gene-mutant/null phenotypic information from the very general at the top to more specific terms in the directed acyclic graph (DAG) by viewing an individual term as a node and its relations to parental terms (allowing for multiple parents) as directed edges. To navigate this hierarchy, we display all parental phenotypic terms to the current phenotypic term of interest ordered by their shortest distances to the current term. Also, only direct children phenotypic terms of the current phenotypic term are listed. Phenotype ontologies we have incorporated are as follows:

Disease Ontology (DO) Ontology (DO)
DO semantically integrates disease and medical vocabularies through extensive cross mapping of DO terms to MeSH, ICD, NCI thesaurus, SNOMED and OMIM.
Human Phenotype (HP) Ontology (HP)
HP captures phenotypic abnormalities that are described in OMIM, along with the corresponding disease-causing genes. It includes three complementary biological concepts: Mode_of_Inheritance (MI), ONset_and_clinical_course (ON), and Phenotypic_Abnormality (PA).
Mouse Phenotype (MP) Ontology (MP)
MP describes phenotypes of the mouse after a specific gene is genetically disrupted. Using it, Mouse Genome Informatics (MGI) provides high-coverate gene-level phenotypes for the mouse.
Worm Phenotype (WP) Ontology (WP)
WP classifies and organizes phenotype descriptions for C. elegans and other nematodes. Using it, WormBase provides primary resource for phenotype annotations for C. elegans.
Yeast Phenotype (YP) Ontology (YP)
Based on YP which is the major contributor to the Ascomycete phenotype ontology, Saccharomyces Genome Database (SGD) provides single mutant phenotypes for every gene in the yeast genome.
Fly Phenotype (FP) Ontology (FP)
FP refers to FlyBase controlled vocabulary. Specifically, a structured controlled vocabulary is used for the annotation of alleles (for their mutagen etc) in FlyBase.
Fly Anatomy (FA) Ontology (FA)
FA is a structured controlled vocabulary of the anatomy of Drosophila melanogaster, used for the description of phenotypes and where a gene is expressed.
Zebrafish Anatomy (ZA) Ontology (ZA)
ZA displays anatomical terms of the zebrafish using standard anatomical nomenclature, together with affected genes.
Xenopus Anatomy (XA) Ontology (XA)
XA represents the lineage of tissues and the timing of development for frogs (Xenopus laevis and Xenopus tropicalis). It is used to annotate Xenopus gene expression patterns and mutant and morphant phenotypes.
Arabidopsis Plant Ontology (AP) Ontology (AP)
As a major contributor to Plant Ontology which describes plant anatomical and morphological structures (AN) and growth and developmental stages (DE), the Arabidopsis Information Resource (TAIR) provides arabidopsis plant ontology annotations for the model higher plant Arabidopsis thaliana.
Enzyme Commission (EC) Ontology (EC)
Each enzyme is allocated a four-digit EC number, the first three digits of which define the reaction catalysed and the fourth of which is a unique identifier (serial number). Each enzyme is also assigned a systematic name that uniquely defines the reaction catalysed.
DrugBank ATC (DB) Ontology (DB)
In the Anatomical Therapeutic Chemical (ATC) classification system, drugs are classified in groups at five different levels according to the organ or system (1st level, anatomical main group) on which they act and their therapeutic (2nd level, therapeutic subgroup), pharmacological (3rd level, pharmacological subgroup) and chemical properties (4th level, chemical subgroup; 5th level, chemical substance). Only drugs in DrugBank are considered.
UniProtKB KeyWords (KW) Ontology (KW)
Keywords in UniProtKB are controlled vocabulary, providing a summary of the entry content and are used to index UniProtKB/Swiss-Prot entries based on 10 categories (the category "Technical term" being excluded here). Each keyword is attributed manually to UniProtKB/Swiss-Prot entries and automatically to UniProtKB/TrEMBL entries (according to specific annotation rules).
UniProtKB UniPathway (UP) Ontology (UP)
UP is a fully manually curated resource for the representation and annotation of metabolic pathways, being used as controlled vocabulary for pathway annotation in UniProtKB.

Structural Domain Phenotype Ontology and its Annotations
Structural Classification of Proteins (SCOP) classifies evolutionary-related domains into Superfamily level and Family level. Using the phenotype ontologies above, we have generated the domain-centric phenotype annotations, and further identified those phenotype terms which are the most informative to annotate SCOP domains. Promisingly, domain-centric phenotypic annotations can serve as an alternative starting point to explore genotype-phenotype relationships. We provide several relevant files for the download, including the annotation and the corresponding ontology for each phenotype ontology.

Structural Domain Disease Ontology (DO) Ontology (SDDO) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2DO.txt, and the corresponding ontology:SDDO.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Human Phenotype (HP) Ontology (SDHP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2HP.txt, and the corresponding ontology:SDHP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Mouse Phenotype (MP) Ontology (SDMP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2MP.txt, and the corresponding ontology:SDMP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Worm Phenotype (WP) Ontology (SDWP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2WP.txt, and the corresponding ontology:SDWP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Yeast Phenotype (YP) Ontology (SDYP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2YP.txt, and the corresponding ontology:SDYP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Fly Phenotype (FP) Ontology (SDFP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2FP.txt, and the corresponding ontology:SDFP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Fly Anatomy (FA) Ontology (SDFA) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2FA.txt, and the corresponding ontology:SDFA.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Zebrafish Anatomy (ZA) Ontology (SDZA) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2ZA.txt, and the corresponding ontology:SDZA.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Xenopus Anatomy (XA) Ontology (SDXA) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2XA.txt, and the corresponding ontology:SDXA.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Arabidopsis Plant Ontology (AP) Ontology (SDAP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2AP.txt, and the corresponding ontology:SDAP.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain Enzyme Commission (EC) Ontology (SDEC) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2EC.txt, and the corresponding ontology:SDEC.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain DrugBank ATC (DB) Ontology (SDDB) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2DB.txt, and the corresponding ontology:SDDB.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain UniProtKB KeyWords (KW) Ontology (SDKW) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2KW.txt, and the corresponding ontology:SDKW.txt) and mysql tables (Domain2PO.sql.gz).
Structural Domain UniProtKB UniPathway (UP) Ontology (SDUP) and its Annotations:
For details, please visit Document: PO annotation for SCOP domains, wherein Data Availability contains parsable flat files (the annotation:Domain2UP.txt, and the corresponding ontology:SDUP.txt) and mysql tables (Domain2PO.sql.gz).

Supra-domain Phenotype Ontology and its Annotations
Although domain-centric annotations hold great promise in describing phenotypic nature of independent domains, most domains themselves may not just work alone. In multi-domain proteins, they may be combined together to form distinct domain architectures. The recombination of the existing domains is considered as one of major driving forces for phenotypic diversificaation. As an extension, we have also generated supra-domain phenotype ontology and its annotations. Compared to domain-centric phenotype ontology and annotations (SCOP domains at the Superfamily level and Family level), this version focuses on supra-domains and individual SCOP domains ONLY at the Superfamily level. Besides, in terms of individual superfamilies, their annotations from the domain-centric version may be different from those from supra-domains version. Depending on your focus, the former should be used for the consideration of both the Superfamily level and Family level, otherwise the latter should be used if you are interested in domain combinations. Also, we provide several relevant files for the download, including the annotation and the corresponding ontology for each phenotype ontology.

Supra-domain Domain Disease Ontology (DO) Ontology (SPDO) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2DO.txt, and the corresponding ontology:SPDO.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Human Phenotype (HP) Ontology (SPHO) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2HP.txt, and the corresponding ontology:SPHO.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Mouse Phenotype (MP) Ontology (SPMP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2MP.txt, and the corresponding ontology:SPMP.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Worm Phenotype (WP) Ontology (SPWP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2WP.txt, and the corresponding ontology:SPWP.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Yeast Phenotype (YP) Ontology (SPYP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2YP.txt, and the corresponding ontology:SPYP.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Fly Phenotype (FP) Ontology (SPFP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2FP.txt, and the corresponding ontology:SPFP.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Fly Anatomy (FA) Ontology (SPFA) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2FA.txt, and the corresponding ontology:SPFA.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Zebrafish Anatomy (ZA) Ontology (SPZA) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2ZA.txt, and the corresponding ontology:SPZA.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Xenopus Anatomy (XA) Ontology (SPXA) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2XA.txt, and the corresponding ontology:SPXA.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Arabidopsis Plant Ontology (AP) Ontology (SPAP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2AP.txt, and the corresponding ontology:SPAP.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain Enzyme Commission (EC) Ontology (SPEC) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2EC.txt, and the corresponding ontology:SPEC.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain DrugBank ATC (DB) Ontology (SPDB) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2DB.txt, and the corresponding ontology:SPDB.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain UniProtKB KeyWords (KW) Ontology (SPKW) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2KW.txt, and the corresponding ontology:SPKW.txt) and mysql tables (SP2PO.sql.gz).
Supra-domain Domain UniProtKB UniPathway (UP) Ontology (SPUP) and its Annotations:
For details, please visit Document: PO annotation for Supra-domains, wherein Data Availability contains parsable flat files (the annotation:SP2UP.txt, and the corresponding ontology:SPUP.txt) and mysql tables (SP2PO.sql.gz).

Jump to [ Top · Phenotype Hierarchy · Superfamily · Family · Supra-domain ]

Root: FA Hierarchy (fly anatomy from FlyBase anatomy ontology)

Shortest distance to current term (+ for parents, - for children) FA term [FA ID] <#Children> (#SF|#FA) #Supra-domain
(Single|Duplex|Triple)

+ + + + + + + + 8: anatomical entity [FBbt:99999999] <4>

+ + + + + + 6: material anatomical entity [FBbt:00007016] <6> (240|245) (239|100|32)

+ + + + + 5: anatomical structure [FBbt:00007001] <6> (240|245) (239|100|32)

+ + + + 4: multicellular structure [FBbt:00100313] <96> (235|240) (235|100|31)

+ + + + 4: organism subdivision [FBbt:00007009] <5> (57|77) (60|34|13)

+ + + 3: organism [FBbt:00000001] <12> (200|210) (201|92|30)

+ + + 3: anterior-posterior subdivision of organism [FBbt:00057001] <4> (56|77) (60|34|13)

+ + 2: tagma [FBbt:00000002] <7> (54|77) (58|34|13)

+ + 2: adult [FBbt:00003004] <46> (66|87) (65|39|16)

+ 1: adult tagma [FBbt:00003005] <3> (41|53) (43|22|11)

+ 1: head [FBbt:00000004] <5> (21|30) (25|15|7)

0: adult head [FBbt:00003007] <30> (18|24) (21|14|7)

- 1: adult brain [FBbt:00003624] <275> (6|5) (8|6|2)

- 1: adult head sensillum [FBbt:00007241] <26> (9|11) (10|7|4)

- 1: skeletal muscle of head [FBbt:00003260] <8> (0|0) (0|0|0)

- 1: adult labial sensory neuron [FBbt:00004029] <7> (0|0) (0|0|0)

- 1: eye [FBbt:00004508] <7> (9|10) (10|6|3)

- 1: head air sac [FBbt:00007134] <5> (0|0) (0|0|0)

- 1: adult external head [FBbt:00004481] <4> (5|5) (5|2|2)

- 1: adult head segment [FBbt:00003009] <2> (9|9) (10|7|2)

- 1: accessory pharyngeal nerve [FBbt:00004024] (0|0) (0|0|0)

- 1: adult frontal nerve [FBbt:00100330] (0|0) (0|0|0)

- 1: adult labial motor neuron [FBbt:00004042] (0|0) (0|0|0)

- 1: adult labial nerve [FBbt:00004028] (0|0) (0|0|0)

- 1: adult labral nerve [FBbt:00100328] (0|0) (0|0|0)

- 1: adult labro-frontal nerve [FBbt:00110653] (0|0) (0|0|0)

- 1: adult maxillary nerve [FBbt:00004043] (0|0) (0|0|0)

- 1: adult maxillary-labial nerve [FBbt:00110654] (0|0) (0|0|0)

- 1: antennal foramen [FBbt:00004487] (0|0) (0|0|0)

- 1: cibarial muscle trachea [FBbt:00003100] (0|0) (0|0|0)

- 1: cibarial pump muscle 11 motor neuron [FBbt:00004023] (0|0) (0|0|0)

- 1: cibarial pump muscle 12 motor neuron [FBbt:00111299] (0|0) (0|0|0)

- 1: cibarial pump muscle 9 motor neuron [FBbt:00111298] (0|0) (0|0|0)

- 1: frontal pulsatile organ [FBbt:00003175] (0|0) (0|0|0)

- 1: labial trachea [FBbt:00003097] (0|0) (0|0|0)

- 1: mandibular nerve [FBbt:00100335] (0|0) (0|0|0)

- 1: maxillary palpus trachea [FBbt:00003098] (0|0) (0|0|0)

- 1: occipital foramen [FBbt:00004499] (0|0) (0|0|0)

- 1: ocellar nerve [FBbt:00003699] (0|0) (0|0|0)

- 1: pericerebral adult fat mass [FBbt:00048409] (0|0) (0|0|0)

- 1: pharyngeal nerve [FBbt:00004021] (0|0) (0|0|0)

- 1: tegumentary nerve [FBbt:00003981] (0|0) (0|0|0)

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Superfamily

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SCOP term	FDR (all)	Annotation (direct or inherited)
Thiamin diphosphate-binding fold (THDP-binding)	0	Direct
Frizzled cysteine-rich domain	0	Direct
SAP domain	0	Direct
Second domain of FERM	0.00007703	Direct
Caspase-like	0.004052	Inherited
SH2 domain	0.03169	Inherited
Cadherin-like	0.03837	Inherited
(Phosphotyrosine protein) phosphatases II	0.06808	Inherited
lambda repressor-like DNA-binding domains	0.1198	Inherited
dsRNA-binding domain-like	0.1574	Inherited
Immunoglobulin	0.2265	Inherited
Fibronectin type III	0.2927	Inherited
EGF/Laminin	0.3028	Inherited
Growth factor receptor domain	0.4258	Inherited
HAD-like	0.672	Inherited
Homeodomain-like	0.7314	Inherited
TPR-like	0.9283	Inherited
HLH, helix-loop-helix DNA-binding domain	1	Inherited

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Family

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SCOP term	FDR (all)	Annotation (direct or inherited)
p120GAP domain-like	0	Direct
Frizzled cysteine-rich domain	0	Direct
Group II chaperonin (CCT, TRIC), intermediate domain	0	Direct
VHS domain	0	Direct
Diacylglycerol kinase-like	0	Direct
SAP domain	0	Direct
Integrin domains	0	Direct
Group II chaperonin (CCT, TRIC), ATPase domain	0	Direct
Group II chaperonin (CCT, TRIC), apical domain	0	Direct
Second domain of FERM	0.0000823	Direct
Third domain of FERM	0.0005986	Direct
Laminin G-like module	0.00145	Inherited
SH2 domain	0.03299	Inherited
Protein kinases, catalytic subunit	0.06083	Inherited
Cadherin	0.06894	Inherited
I set domains	0.1034	Inherited
POU-specific domain	0.1618	Inherited
Extracellular domain of cell surface receptors	0.2324	Inherited
Fibronectin type III	0.3148	Inherited
EGF-type module	0.3532	Inherited
Growth factor receptor domain	0.3634	Inherited
BTB/POZ domain	0.6219	Inherited
Homeodomain	0.8872	Inherited
HLH, helix-loop-helix DNA-binding domain	1	Inherited

Plot distribution on phylogenetic tree for Superfamily and/or Family domains annotated by this phenotype term

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Jump to [ Top · Phenotype Hierarchy · Superfamily · Family · Supra-domain ]

Supra-domain (including individual superfamily)

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Supra-domain (Single)	FDR (all)	Annotation (direct or inherited)
SAP domain	0	Direct
Frizzled cysteine-rich domain	0	Direct
Thiamin diphosphate-binding fold (THDP-binding)	0	Direct
Second domain of FERM	0.00005645	Direct
Caspase-like	0.002922	Inherited
SH2 domain	0.0248	Inherited
Cadherin-like	0.03019	Inherited
Protein kinase-like (PK-like)	0.04345	Inherited
Plexin repeat	0.05395	Inherited
(Phosphotyrosine protein) phosphatases II	0.05543	Inherited
lambda repressor-like DNA-binding domains	0.1011	Inherited
dsRNA-binding domain-like	0.1348	Inherited
Snake toxin-like	0.149	Inherited
Immunoglobulin	0.1988	Inherited
Fibronectin type III	0.2619	Inherited
EGF/Laminin	0.2722	Inherited
Growth factor receptor domain	0.3933	Inherited
HAD-like	0.6453	Inherited
Homeodomain-like	0.7084	Inherited
TPR-like	0.9185	Inherited
HLH, helix-loop-helix DNA-binding domain	1	Inherited

(show details)

Supra-domain (Duplex) in N- to C-terminal order	FDR (all)	Annotation (direct or inherited)
55550,50044 55550 - SH2 domain 50044 - SH3-domain	0	Direct
50044,55550 50044 - SH3-domain 55550 - SH2 domain	0	Direct
54236,47031 54236 - Ubiquitin-like 47031 - Second domain of FERM	0.0004303	Direct
49899,49899 49899 - Concanavalin A-like lectins/glucanases 49899 - Concanavalin A-like lectins/glucanases	0.0004303	Direct
47031,50729 47031 - Second domain of FERM 50729 - PH domain-like	0.0004303	Direct
49313,49313 49313 - Cadherin-like 49313 - Cadherin-like	0.05788	Inherited
48726,48726 48726 - Immunoglobulin 48726 - Immunoglobulin	0.08604	Inherited
46966,46966 46966 - Spectrin repeat 46966 - Spectrin repeat	0.1011	Inherited
47413,46689 47413 - lambda repressor-like DNA-binding domains 46689 - Homeodomain-like	0.1011	Inherited
51045,51045 51045 - WW domain 51045 - WW domain	0.1011	Inherited
48726,49265 48726 - Immunoglobulin 49265 - Fibronectin type III	0.2015	Inherited
57196,57184 57196 - EGF/Laminin 57184 - Growth factor receptor domain	0.2184	Inherited
57196,57196 57196 - EGF/Laminin 57196 - EGF/Laminin	0.3178	Inherited
49265,49265 49265 - Fibronectin type III 49265 - Fibronectin type III	0.4445	Inherited

(show details)

Supra-domain (Triple) in N- to C-terminal order	FDR (all)	Annotation (direct or inherited)
57196,49899,49899 57196 - EGF/Laminin 49899 - Concanavalin A-like lectins/glucanases 49899 - Concanavalin A-like lectins/glucanases	0	Direct
49899,49899,49899 49899 - Concanavalin A-like lectins/glucanases 49899 - Concanavalin A-like lectins/glucanases 49899 - Concanavalin A-like lectins/glucanases	0	Direct
54236,47031,50729 54236 - Ubiquitin-like 47031 - Second domain of FERM 50729 - PH domain-like	0.002922	Inherited
49313,49313,49313 49313 - Cadherin-like 49313 - Cadherin-like 49313 - Cadherin-like	0.05788	Inherited
57196,57196,57196 57196 - EGF/Laminin 57196 - EGF/Laminin 57196 - EGF/Laminin	0.09806	Inherited
48726,48726,48726 48726 - Immunoglobulin 48726 - Immunoglobulin 48726 - Immunoglobulin	0.1036	Inherited
48726,48726,49265 48726 - Immunoglobulin 48726 - Immunoglobulin 49265 - Fibronectin type III	0.2552	Inherited

Plot distribution on phylogenetic tree for Supra-domains (Single/Individual) annotated by this phenotype term

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Fly Anatomy (FA): adult head

Superfamily

Family

Supra-domain (including individual superfamily)